Petiole borer (Lepidoptera: Tortricidae) in Aesculus

I found larvae tunneling in petioles of new leaves in spring. Their activity caused the leaflets to shrivel and the leaf above the tunneled area to droop conspicuously. The petiole sometimes discolored and/or became slightly distended, and a hole was sometimes visible in the petiole wall, through which the larva evidently expelled frass. Inhabited leaves eventually fell to the ground, and larvae completed their development by exiting the petiole and feeding on the leaflets, which they tied together with silk, further fortifying the shelter already established by the complex infolded architecture of the shriveled leaf blade tissue. Adults emerged in June. S. Nanz and M. Sabourin reviewed photos of the adults (Nanz 2019), and Eiseman's (2014) images of larval damage, and also an adult he reared that was examined by J. Brown to confirm its identity, provided a helpful point of comparison.